1 Key Laboratory of Computational Biology, CAS-MPG Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Shanghai, China. liyang@picb.ac.cn linglingchen@sibcb.ac.cn.
2 State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, CAS Center for Excellence in Molecular Cell Science, Shanghai Institute of Biochemistry and Cell Biology, Chinese Academy of Sciences, Shanghai, China. liyang@picb.ac.cn linglingchen@sibcb.ac.cn.
3 School of Life Science and Technology, ShanghaiTech University, Shanghai, China.