1 Department of Ecology and Genetics, Evolutionary Biology Centre, Uppsala University, Norbyvägen 18D, Uppsala 75236 Sweden laura.parducci@ebc.uu.se.
2 Geography & Sustainable Development, University of St Andrews, St Andrews Fife KY16 9AL, Scotland, UK and Marine Laboratory, Queen's University Belfast, Portaferry, BT22 1LS, Northern Ireland, UK.