1 Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, 100101, China.
2 Shanghai Key Laboratory for Contemporary Applied Mathematics, School of Mathematical Sciences, Fudan University, Shanghai, 200433, China.
3 School of Life Sciences, Nanchang University, Nanchang, 330031, China.
4 College of Life Sciences, University of the Chinese Academy of Sciences, Beijing, 100049, China.
5 Shanghai Key Laboratory for Contemporary Applied Mathematics, School of Mathematical Sciences, Fudan University, Shanghai, 200433, China. zmwu@fudan.edu.cn.
6 Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, 100101, China. weifw@ioz.ac.cn.