1 Guangdong Provincial Key Laboratory for Plant Epigenetics, College of Life Sciences and Oceanography, Shenzhen University, Shenzhen, Guangdong 518060, P.R. China
2 Key Laboratory of Optoelectronic Devices and Systems of Ministry of Education and Guangdong Province, College of Optoelectronic Engineering, Shenzhen University, Shenzhen, Guangdong 518060, P.R. China
3 Department of Molecular, Cellular and Developmental Biology, Yale University, New Haven, CT 06520, USA
4 Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06520, USA