1 Molecular Ecology & Fisheries Genetics Laboratory, School of Biological Sciences, Bangor University, Deiniol Road, Bangor, LL57 2UW, U.K.
2 School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA, 98195-5020, U.S.A.
3 European Commission, Joint Research Centre (JRC), Directorate D - Sustainable Resources, Water and Marine Resources, Via E. Fermi 2749, I-21027, Ispra, VA, Italy.