1 Department of Geoscience, James Cook University, Townsville, Qld 4811, Australia; Evolutionary Studies Institute, DST/NFR Centre of Excellence in Palaeosciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa. Electronic address: paul.dirks@jcu.edu.au.
2 Evolutionary Studies Institute, DST/NFR Centre of Excellence in Palaeosciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa.
3 Evolutionary Studies Institute, DST/NFR Centre of Excellence in Palaeosciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa; Department of Anthropology, University of Wisconsin-Madison, Madison, WI 53593, USA.
4 Evolutionary Studies Institute, DST/NFR Centre of Excellence in Palaeosciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa; School of Anatomical Sciences, University of the Witwatersrand Medical School, Johannesburg 2193, South Africa.
5 Department of Geoscience, James Cook University, Townsville, Qld 4811, Australia; Evolutionary Studies Institute, DST/NFR Centre of Excellence in Palaeosciences, University of the Witwatersrand, Private Bag 3, Wits 2050, South Africa.