1 Department of Neurobiology, University of Massachusetts Medical School, Worcester, MA 01605; michael.francis@umassmed.edu marc.freeman@umassmed.edu.
2 Department of Neurobiology, University of Massachusetts Medical School, Worcester, MA 01605; Howard Hughes Medical Institute, University of Massachusetts Medical School, Worcester, MA 01605 michael.francis@umassmed.edu marc.freeman@umassmed.edu.