1 Stazione Zoologica Anton Dohrn, Villa Comunale, 80121 Napoli, Italy.
2 Department of Biology and Evolution of Marine Organisms, Stazione Zoologica Anton Dohrn, Villa Comunale, 80121 Napoli, Italy.
3 National Research Council, Via Pietro Castellino 111, 80131 Napoli, Italy; Roma 3 University, Department of Biology, Viale Marconi 446, 00146 Roma, Italy.