1 Research Department of Cell and Developmental Biology, UCL, Gower Street, London, WC1E 6BT, UK.
2 Department of Cell and Molecular Biology, University of A Coruña, 15008, A Coruña, Spain.
3 Sars Centre for Marine Molecular Biology, University of Bergen, 5008, Bergen, Norway.
4 Developmental Neurobiology and Genomics, Brain and Mind Research Institute, Sydney Medical School, University of Sydney, Camperdown, NSW, 2050, Australia.
5 Research Department of Cell and Developmental Biology, UCL, Gower Street, London, WC1E 6BT, UK. jon.clarke@kcl.ac.uk.
6 MRC Centre for Developmental Neurobiology, King's College London, Guy's Hospital Campus, London, SE1 1UL, UK. jon.clarke@kcl.ac.uk.