1 Laboratoire d'Ecologie, Systématique, et Evolution, Unité Mixte de Recherche (UMR) 8079, Université Paris Sud, Orsay, France; Department of Ecology and Evolutionary Biology and Center for Tropical Research, and the Institute of the Environment and Sustainability, University of California Los Angeles, Los Angeles, CA 90095, USA.
2 Santa Fe Institute, 1399 Hyde Park Road, Santa Fe, USA.
3 Institut Pluridisciplinaire Hubert Curien, CNRS UMR 7178, Université de Strasbourg, France.
4 Department of Zoology, University of Oxford, Oxford OX1 3PS, UK.
5 Laboratoire Evolution et Diversité Biologique, CNRS UMR 5174, Université Paul Sabatier, Toulouse, France.
6 Santa Fe Institute, 1399 Hyde Park Road, Santa Fe, USA; Institut des Sciences de l'Evolution, CNRS UMR 5554, Université Montpellier II, Montpellier, France. Electronic address: mhochber@univ-montp2.fr.