1 Department of Biology, Research Group Ecosystem Management, University of Antwerp, Wilrijk, Belgium.
2 Department of Geology and Mineralogy, Royal Museum for Central Africa, Tervuren, Belgium.
3 Department of Earth and Environmental Sciences, Catholic University of Leuven, Heverlee, Belgium.
4 Department of Geology, Lund University, Lund, Sweden.
5 1] Department BIOSystem Engineering, University of Liège, Gembloux, Belgium [2] Earth and Life Institute, Université catholique de Louvain, Louvain-la-Neuve, Belgium.