1 School of Biological Sciences, University of Wollongong, Wollongong, NSW, Australia, 2522; Centre for Evolutionary Biology, School of Animal Biology, University of Western Australia, Crawley, WA, Australia, 6008; Centre of Excellence in Biological Interactions, University of Jyväskylä, Jyväskylä, Finland. Electronic address: kateu@uow.edu.au.
2 Evolutionary Biology, Zoological Institute, University of Basel, Vesalgasse 1, CH-4051 Basel, Switzerland.
3 Centre of Excellence in Biological Interactions, University of Jyväskylä, Jyväskylä, Finland.