1 Department of Geography and Environmental Science, University of Reading, Whiteknights, Reading RG6 6AB, United Kingdom; School of GeoSciences, University of Edinburgh, Edinburgh EH8 9XP, United Kingdom; j.f.carson@reading.ac.uk.
2 School of GeoSciences, University of Edinburgh, Edinburgh EH8 9XP, United Kingdom;
3 Department of Geography and Environmental Science, University of Reading, Whiteknights, Reading RG6 6AB, United Kingdom;
4 Department of Archaeology, University of Exeter, Exeter EX4 4QE, United Kingdom;
6 Herbario del Oriente Boliviano, Museo de Historia Natural 'Noel Kempff Mercado,' Universidad Autónoma Gabriel René Moreno, Casilla 2489, Santa Cruz, Bolivia.