1 Sea Mammal Research Unit, University of St Andrews, St Andrews, Fife, KY16 8LB, UK; Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St Andrews, Fife, KY16 9LZ, UK. Electronic address: dr60@st-and.ac.uk.
2 IMARES, Wageningen University and Research Centre, Landsdiep 4, 1797 SZ 't Horntje -Texel, The Netherlands.
3 Sea Mammal Research Unit, University of St Andrews, St Andrews, Fife, KY16 8LB, UK.
4 IMARES, Wageningen University and Research Centre, Landsdiep 4, 1797 SZ 't Horntje -Texel, The Netherlands; Aquatic Ecology and Water Quality Management, Wageningen University, P.O. Box 47, 6700 AA Wageningen, The Netherlands.
5 National Marine Mammal Laboratory, Alaska Fisheries Science Center, NOAA-NMFS, 7600 Sand Point Way NE, Seattle, Washington 98115, USA.
6 Institute of Biodiversity Animal Health and Comparative Medicine, Graham Kerr Building, University of Glasgow, Glasgow, G12 8QQ, UK.