1 University of Montpellier II, Place Eugène Bataillon, 34095 Montpellier Cedex 5, France.
2 1] University of Montpellier II, Place Eugène Bataillon, 34095 Montpellier Cedex 5, France [2] CNRS, Institute of Evolutionary Sciences, CC 065, Place Eugène Bataillon, Montpellier 34095, France.