1 Centre of Excellence in Biological Interactions and Division of Evolution Ecology and Genetics, Research School of Biology, The Australian National University, Canberra, ACT 0200, Australia. Electronic address: jussi.lehtonen@iki.fi.
2 Centre of Excellence in Biological Interactions and Division of Evolution Ecology and Genetics, Research School of Biology, The Australian National University, Canberra, ACT 0200, Australia.