In the ectomycorrhizal symbiosis between fungi and trees, the fungus completely ensheaths the tree roots and takes over water and mineral nutrient supply, while the plant supplies photosynthate. Recent work has focussed on gene expression in the two partners, on the effects of global change and nitrogen deposition rate on the symbiosis, and on the role of mycorrhizal fungi in connecting individual plants to form a 'wood-wide web'.