In a population consisting of 227 test-crosses of Zhenshan 97A x (Zhenshan 97B x Milyang 46) F6, a linkage map of 115 RFLP markers was constructed and employed to detect QTL for rice CMS-WA (wild abortive cytoplasmic male sterility) fertility restoration. One QTL having major effect, qRf-10, and 3 QTL having minor effects, qRf-1, qRf-7 and qRf-11, were detected. The four genes mainly acted additively with each other, but epistasis was also evident. Two minor QTL, qRf-1 and qRf-11, were significantly interacted with the major gene qRf-10. It was also found that interaction effects among minor QTL varied greatly depending on whether qRf-10 was present. Two other QTL, qSF-1 and qSF-7, were detected, which did not display any effects on fertility restoration. Instead, they showed effects for increasing SF when qRf-10 was present.