The First Representative of the Roachoid Family Spiloblattinidae (Insecta, Dictyoptera) from the Late Pennsylvanian of the Iberian Peninsula

André Nel; Artai A Santos; Antonio Hernández-Orúe; Torsten Wappler; José B Diez; Enrique Peñalver

doi:10.3390/insects13090828

The First Representative of the Roachoid Family Spiloblattinidae (Insecta, Dictyoptera) from the Late Pennsylvanian of the Iberian Peninsula

Insects. 2022 Sep 12;13(9):828. doi: 10.3390/insects13090828.

Authors

André Nel¹, Artai A Santos^{2

3}, Antonio Hernández-Orúe⁴, Torsten Wappler^{5

6}, José B Diez^{2

3}, Enrique Peñalver⁷

Affiliations

¹ Institut Systématique Evolution Biodiversité (ISYEB), Museum National d'Histoire Naturelle, CNRS, Sorbonne Université, Université des Antilles, EPHE, 57 Rue Cuvier, CP 50, 75005 Paris, France.
² Departamento de Xeociencias Mariñas e Ordenación do Territorio, Facultade de Ciencias do Mar, Universidade de Vigo, 36310 Vigo, Spain.
³ Centro de Investigación Mariña, Universidade de Vigo (CIM-UVIGO), 36310 Vigo, Spain.
⁴ Departamento de Geología, Universidad del País Vasco/Euskal Herriko Unibertsitatea (UPV/EHU), Part 644, 48080 Bilbao, Spain.
⁵ Department of Natural History, Hessisches Landesmuseum Darmstadt, Friedensplatz 1, 64283 Darmstadt, Germany.
⁶ Department of Palaeontology, Institute of Geosciences, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany.
⁷ Instituto Geológico y Minero de España (IGME), CSIC, C/Cirilo Amorós 42, 46004 Valencia, Spain.

Abstract

Sysciophlebia 'sp. form Villablino', the first Iberian representative of the Palaeozoic-Early Mesozoic family Spiloblattinidae, is described and illustrated. Its forewing colour pattern is strongly similar to those of the Gzhelian-early-middle Asselian species Sysciophlebia euglyptica, Sysciophlebia ilfeldensis, Sysciophlebia rubida, and 'Sysciophlebia sp. form KBQ', supporting the currently proposed Gzhelian age for its type locality. It supports the use of the representatives of the Spiloblattinidae for stratigraphic purposes. The diagnoses and limits of the families Subioblattidae, Phyloblattidae, Compsoblattidae, Spiloblattinidae, and of the spiloblattinid genera are discussed.

Keywords: Carboniferous; Gzhelian; Holopandictyoptera; Pennsylvanian; Polyneoptera; Sysciophlebia; roachoid families and genera; stratigraphic markers; taxonomy.

Grants and funding

This research received no external funding.