1 Key Laboratory of Watershed Geographic Sciences, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, China.
2 Biology Department, San Diego State University, San Diego, CA 92182, USA.
3 School of the Environment, Yale University, New Haven, CT 06511, USA.
4 Key Laboratory of Watershed Geographic Sciences, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences, Nanjing 210008, China; University of Chinese Academy of Sciences, Nanjing (UCASNJ), Nanjing 211135, China. Electronic address: htduan@niglas.ac.cn.