1 Federal Scientific Center of the East Asia Terrestrial Biodiversity; Far East Branch of the Russian Academy of Sciences; 100 let Vladivostoku 159; 690022 Vladivostok; Russia. makarchenko@biosoil.ru.
2 A.N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences; Leninskij prosp. 33; 119071 Moscow; Russia. triops@yandex.ru.