1 Department of Fish, Wildlife and Conservation Biology, Colorado State University, 80523, Fort Collins, CO, USA. g.wittemyer@colostate.edu.
2 Quantitative Disease Ecology and Conservation (QDEC) Lab, Department of Geography and the Emerging Pathogens Institute, University of Florida, 32611, Gainesville, FL, USA.
3 School of Life Sciences, University of KwaZulu-Natal, Durban, South Africa.