1 The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, School of Life Sciences, Shandong University, Qingdao 266237, China.
2 The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, School of Life Sciences, Shandong University, Qingdao 266237, China. Electronic address: jiaqi.sun@sdu.edu.cn.
3 Department of Biology, McGill University, Montreal, QC H3A 1B1, Canada. Electronic address: hugo.zheng@mcgill.ca.