1 NIHR Nottingham Biomedical Research Centre, University of Nottingham, Nottingham, UK tanya.monaghan@nottingham.ac.uk.
2 Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham, UK.
3 Department of Biosciences, Centre for Health, Ageing and Understanding Disease (CHAUD), School of Science and Technology, Nottingham Trent University, Nottingham, UK.
4 School of Medicine, Biodiscovery Institute, University of Nottingham, Nottingham, UK.
5 Division of Molecular Therapeutics & Formulation, School of Pharmacy, University of Nottingham, Nottingham, UK.
6 School of Veterinary Medicine and Science, University of Nottingham, Sutton Bonington, UK.
7 MRC Versus Arthritis Centre for Musculoskeletal Ageing Research, Nottingham, UK.
8 Division of Gastroenterology, Department of Medicine, University of Alberta Hospital, Edmonton, Alberta, Canada.
9 John van Geest Cancer Research Centre, School of Science & Technology, Nottingham Trent University, Nottingham, UK.