1 Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, P. R. China. The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, P. R. China. The Provincial Key Laboratory for Agricultural Pest Man-agement of Mountainous Region, Guiyang 550025, Guizhou, P. R. China.. maoshuwuyouzhi@163.com.
2 Laboratório de Sistemática e Biologia de Coleoptera, Departamento de Biologia Animal, Universidade Federal de Viçosa, 36570-900, Viçosa, Minas Gerais, Brazil. . ciidae@gmail.com.
3 Zhengzhou Foreign Language School-New Fengyang Campus, 66 Cuizhu Street, Zhengzhou, 450001, P. R. China. . 3137068712@qq.com.
4 Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, P. R. China. 2The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, P. R. China. 3The Provincial Key Laboratory for Agricultural Pest Man-agement of Mountainous Region, Guiyang 550025, Guizhou, P. R. China.. chenxs3218@163.com.