1 School of Mathematics, College of Science and Engineering, University of Edinburgh, Edinburgh, UK. Electronic address: Jamie.Burke@ed.ac.uk.
2 British Heart Foundation Centre for Cardiovascular Science, University of Edinburgh, Edinburgh, UK.
3 Princess Alexandra Eye Pavilion, NHS Lothian, Edinburgh, UK; Centre for Clinical Brain Sciences, School of Clinical Sciences, University of Edinburgh, Edinburgh, UK.
4 Department of Eye & Vision Sciences, University of Liverpool, Liverpool, UK; Malawi-Liverpool-Wellcome Trust, Blantyre, Malawi.
5 Department of Eye & Vision Sciences, University of Liverpool, Liverpool, UK; St Paul's Eye Unit, Royal Liverpool University Hospital, Liverpool, UK.
6 Centre for Inflammation Research, The Queen's Medical Research Institute, University of Edinburgh, Edinburgh, UK; Institute for Adaptive and Neural Computation, School of Informatics, University of Edinburgh, Edinburgh, UK.