1 Key Laboratory of Freshwater Fish Reproduction and Development (Ministry of Education), Southwest University School of Life Sciences, Chongqing 400715, China.
2 College of Fisheries, Southwest University, Rongchang, Chongqing 402460, China.
3 Academy of Plateau Science and Sustainability, Qinghai Normal University, Xining, Qinghai 810008, China. E-mail: pzg@swu.edu.cn.
This work was supported by the National Natural Science Foundation of China (31872204) and Fundamental Research Funds for the Central Universities (SWU120049)