1 Institute of Oceanography, College of Geography and Oceanography, Minjiang University, Fuzhou, 350108, China.
2 Fujian Provincial Key Laboratory on Conservation and Sustainable Utilization of Marine Biodiversity, Minjiang University, Fuzhou, 350108, China.
3 Technology Innovation Center for Monitoring and Restoration Engineering of Ecological Fragile Zone in Southeast China, Ministry of Natural Resources, Fuzhou, 350108, China.