1 Key Laboratory of Marine Genetics and Breeding (Ocean University of China), Ministry of Education, Qingdao, China.
2 Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China.
3 Function Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China.
4 CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China.