1 Jilin Provincial Key Laboratory of Grassland Farming, Northeast Institute of Geography and Agoecology, Chinese Academy of Sciences, Changchun, China.
2 China Agricultural University, Beijing, China.
3 Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, China.
4 NHC Key Laboratory of Human Disease Comparative Medicine, Peking Union Medical College, Institute of Laboratory Animal Sciences, Chinese Academy of Medical Sciences and Comparative Medicine Center, Beijing, China.
5 Baylor College of Medicine, Houston, TX, United States.