1 Quantitative Plant Ecology and Biodiversity Research Lab, Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran.
2 Quantitative Plant Ecology and Biodiversity Research Lab, Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran. hejtehadi@um.ac.ir.
3 Department of Range and Watershed Management, Faculty of Natural Resources and Environment, Ferdowsi University of Mashhad, Mashhad, Iran. mjankju@um.ac.ir.
4 Botanical Garden of Nowshahr, Research Institute of Forests and Rangelands, Agricultural Research, Education and Extension Organization, AREEO, Tehran, Iran.
5 Department of Range and Watershed Management, Faculty of Natural Resources and Environment, Ferdowsi University of Mashhad, Mashhad, Iran.