1 IRD, CNRS, University Paris-Saclay, UMR Evolution, Génome, Comportement et Ecologie (EGCE), Institut Diversité Ecologie et Evolution du Vivant (IDEEV), 12 Route 128, 91198 Gif‑sur‑Yvette, France.
2 Botswana International University of Science and Technology, Palapye, Botswana; Department of Zoology and Entomology, Rhodes University, Makhanda 6140, South Africa. Electronic address: nyamukondiwac@biust.ac.bw.