1 Biodiversity Division, National Institute for Environmental Studies, Tsukuba, Ibaraki, 305-8506, Japan. nozaki@bs.s.u-tokyo.ac.jp.
2 Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, 113-0033, Japan. nozaki@bs.s.u-tokyo.ac.jp.
3 Biodiversity Division, National Institute for Environmental Studies, Tsukuba, Ibaraki, 305-8506, Japan.
4 Global Environmental Forum, Inarimae, Ibaraki, Tsukuba-shi, 305-0061, Japan.
5 Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, 305-8572, Japan.