1 Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Economic Animals & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Zhanjiang, China.
2 Guangdong Provincial Key Laboratory for Healthy and Safe Aquaculture, Guangdong Provincial Engineering Technology Research Center for Environmentally-Friendly Aquaculture, School of Life Sciences, South China Normal University, Guangzhou, China.
3 Universitetet i Tromsø (UiT) The Arctic University of Norway, Tromsø, Norway.
4 Faculty of Marine Science and Technology, Fukui Prefectural University, Obama, Japan.
5 College of Bioresource Sciences, Nihon University, Fujisawa, Japan.