1 Department of Biological Sciences, Korea Advanced Institute of Science and Technology, 291 Daehak-ro, Yuseong-gu, Daejeon, 34141, South Korea.
2 Korea Crop Environment Research Division, National Institute of Crop Science, 126 Suin-ro, Seodun-dong, Gwonseon-gu, Suwon, Gyeonggi-do, 16429, South Korea.
3 Crop Foundation Research Division, National Institute of Crop Science, 181 Hyeoksin-ro, Iseo-myeon, Jeollabuk-do, 55365, Wanju-gun, South Korea.
4 Department of Biological Sciences, Korea Advanced Institute of Science and Technology, 291 Daehak-ro, Yuseong-gu, Daejeon, 34141, South Korea. sgkim1@kaist.ac.kr.