1 Key Laboratory of Horticultural Plant Biology (Ministry of Education), Huazhong Agricultural University, Wuhan, China.
2 Hubei Hongshan Laboratory, Wuhan, China.
3 The New Zealand Institute for Plant and Food Research Ltd, Auckland, New Zealand.
4 Department of Cell Biology and Molecular Genetics, University of Maryland, College Park, MD, United States.
5 Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, The Innovative Academy of Seed Design, Chinese Academy of Sciences, Wuhan, China.