1 Institute of Systematic Botany and Ecology, Ulm University, Albert-Einstein-Allee 11, 89081, Ulm, Germany.
2 College of Life and Environmental Sciences, University of Exeter, Exeter, EX4 4RJ, UK.
3 Department of Biological Science, California State University Fullerton, 800 N. State College Boulevard, Fullerton, CA, 92831, USA.
4 Department of Integrative Biology and Biodiversity Research, Institute of Botany, University of Natural Resources and Life Sciences, Gregor-Mendel-Straße 33, 1180, Vienna, Austria.