1 Institute for Biological Problems of the North RAS, Portovaya Str.18, Magadan, 685000, Russia. Department of Zoology Entomology, University of the Free State, Bloemfontein 9300, South Africa.. yurmar@mail.ru.
2 National Museums Scotland Chambers Street Edinburgh EH1 1JF, UK. v.blagoderov@nms.ac.uk.