1 Co-Innovation Center for Sustainable Forestry in Southern China & Key Laboratory of National Forestry and Grassland Administration on Subtropical Forest Biodiversity Conservation, College of Biology and the Environment, Nanjing Forestry University, Nanjing 210037, China.
2 Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China.
3 Missouri Botanical Garden, St. Louis, MO 63110, USA.
4 Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China.