1 Centre for Biodiversity & Environment Research, University College London, Gower Street, London, WC1E 6BT, UK.
2 Dipartimento di Biologia, Università degli Studi di Firenze, Via Madonna del Piano, Sesto Fiorentino, Firenze, 50019, Italy.
3 Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, Turin, 10123, Italy.
4 Department of Evolutionary Biology and Ecology, Institute of Biology I (Zoology), University of Freiburg, Hauptstraße 1, Freiburg (Brsg.), 79104, Germany.