1 Department of Biology, University of Pennsylvania, Philadelphia, PA, USA.
2 State Key Laboratory of Medical Neurobiology, MOE Frontiers Center for Brain Science, School of Life Sciences, Fudan University, Shanghai, China.
3 Department of Neurology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
4 School of Life Sciences, IDG/McGovern Institute for Brain Research, Peking University, Beijing, China.
5 Center for Neurodegenerative Disease Research, Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
6 Department of Neurology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA. chenplot@pennmedicine.upenn.edu.
7 Center for Neurodegenerative Disease Research, Department of Pathology and Laboratory Medicine, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA. kelvincl@pennmedicine.upenn.edu.
8 Department of Biology, University of Pennsylvania, Philadelphia, PA, USA. dren@sas.upenn.edu.