1 Sue and Bill Gross Stem Cell Research Center and Departments of Developmental and Cell Biology and Biological Chemistry, University of California, Irvine, CA 92697, USA. pdonovan@uci.edu zaret@pennmedicine.upenn.edu.
2 Institute for Regenerative Medicine and Department of Cell and Developmental Biology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA. pdonovan@uci.edu zaret@pennmedicine.upenn.edu.