1 State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, 100093, Beijing, China. gesong@ibcas.ac.cn.
2 University of Chinese Academy of Sciences, 100049, Beijing, China. gesong@ibcas.ac.cn.
3 State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, 100093, Beijing, China.
4 University of Chinese Academy of Sciences, 100049, Beijing, China.