Photosynthesis is the gateway of the Sun's energy into the biosphere and the source of the ozone layer; thus it is both provider and protector of life as we know it. Despite its pivotal role we know surprisingly little about the genetic basis of variation in photosynthesis and the selective pressures giving rise to or maintaining this variation. In this review, I will briefly summarise our current knowledge of intraspecific and interspecific variation in photosynthesis to understand the main selective constraints on photosynthesis and what this means for the future of nature and agriculture in a changing world.
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