1 Institute for Frontier Life and Medical Sciences, Kyoto University, Shogoin-Kawahara, Sakyo-Ku, Kyoto, 606-8507, Japan. Electronic address: rkageyam@infront.kyoto-u.ac.jp.
2 Keio University School of Medicine, Shinanomachi, Shinjuku-ku, Tokyo, 160-8582, Japan.