1 Centre for Sustainable Aquatic Ecosystems, Harry Butler Institute, Murdoch University, Murdoch, WA6150, Australia. Electronic address: M.Keulen@murdoch.edu.au.
2 Natural Resources and Sustainable Development, Department of Earth Sciences, Uppsala University, Villavägen 16, 75 236, Uppsala, Sweden.
3 Sustainable Places Research Institute, Cardiff University, 33 Park Place, Cardiff, CF10 3BA, UK.