1 Department of Pharmacology, University of Oxford, Oxford OX1 3QT, United Kingdom and matthew.buchan@chch.ox.ac.uk james.rowland@ccc.ox.ac.uk.
2 Department of Physiology, Anatomy & Genetics, University of Oxford, Oxford OX1 3QX, United Kingdom matthew.buchan@chch.ox.ac.uk james.rowland@ccc.ox.ac.uk.