1 Department of Genetics and Center for Molecular Studies in Digestive and Liver Diseases, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
2 Department of Molecular Cell Biology, Weizmann Institute of Science, Rehovot, Israel.
3 Department of Genetics and Center for Molecular Studies in Digestive and Liver Diseases, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA. kaestner@pennmedicine.upenn.edu.