1 Hokkaido Agriculture Research Centre, National Agriculture and Food Research Organization Toyohira-ku, Sapporo, 062-8555, Japan.
2 Biotechnology Development Laboratories, KANEKA CORPORATION, Takasago, 6768688, Japan.
3 Hokkaido Agriculture Research Centre, National Agriculture and Food Research Organization Toyohira-ku, Sapporo, 062-8555, Japan. rzi@affrc.go.jp.
4 Institute of Agrobiological Sciences, National Agriculture and Food Research Organization, 2-1-2 Kannondai, Tsukuba, 305-8602, Japan. rzi@affrc.go.jp.