1 The School of Energy, Geoscience, Infrastructure and Society, Institute of Life and Earth Sciences, Heriot-Watt University , Edinburgh, EH14 4AS U.K.
2 The Center for Environmental Biotechnology, The University of Tennessee , Knoxville, Tennessee 37996 United States.
3 Department of Forestry Wildlife and Fisheries, The University of Tennessee , Knoxville, Tennessee 37996, United States.
4 Hospital for Small Animals, Royal (Dick) School of Veterinary Studies and the Roslin Institute, University of Edinburgh , Midlothian, EH25 9RG U.K.